即使是再微不足道的小人物，也有怀揣梦想的权力，哪怕他只是一只生活在阴沟里的老鼠呢……而他的梦想，是成为法国五星级饭店厨房的掌勺。
由于在嗅觉方面有着无与伦比的天赋，雷米的一生都浸透在“厨师” 的光辉理想之中，并且努力地在朝这个方向艰难地迈进，丝毫不去理会摆在自己面前的事实：厨师是这个世界上对老鼠最怀有病态恐惧的职业。至少他的家人都还算正常，而且对于雷米的异想天开皆嗤之以鼻，快乐而满足地过着老鼠都在过的与丢弃不要的东西堆为伴的生活。这个时候的雷米已经有点走火入魔了，凡是看到能吃的东西，他都会情不自禁地想象：到底是火烧、还是嫩煎好呢……
你还别不信，有的时候梦想和现实，真的只有一步之遥。雷米在一个偶然的机会里，竟然搬到一家法国餐馆的下水道里安家。这还不算什么，最神的是，这家餐馆的创始人，恰恰就是雷米毕生的偶像--法国名厨奥古斯汀·古斯特，他曾说过的那句“人人都能当厨师”早就被雷米奉为金玉良言。可是雷米也有自己的麻烦，因为他不能让自己在厨房中被发现，否则就会引起惊天动地的可怕混乱。
就在雷米饱受折磨准备放弃的时候，却发现餐馆厨房里有一个倒霉的学徒林奎尼，因为生性害羞而遭排斥的他，在厨艺上更是没什么天赋，即将面对被解雇的命运。于是乎，同被逼上绝境的一人一鼠，竟然结成了一个不可思议的同盟：林奎尼以人的身份在前台“表演”，雷米则奉献了他那有创造力的大脑，在幕后进行操纵--没想到，他们竟然共同获得了不可思议的成功。林奎尼在雷米的帮助下，成为了整个法国饮食业的“天才厨师”，他们撼动的，可不仅仅是巴黎，还包括整个世界的价值观。
A rat named Remy dreams of becoming a great French chef despite his family's wishes and the obvious problem of being a rat in a decidedly rodent-phobic profession. When fate places Remy in the sewers of Paris, he finds himself ideally situated beneath a restaurant made famous by his culinary hero, Auguste Gusteau. Despite the apparent dangers of being an unlikely - and certainly unwanted - visitor in the kitchen of a fine French restaurant, Remy's passion for cooking soon sets into motion a hilarious and exciting rat race that turns the culinary world of Paris upside down.
Remy, a provincial rat with a wonderful sense of smell, hates garbage and risks death to enter a human kitchen where he discovers real food and the cooking of five-star chef, Anton Gusteau, author of "Anyone Can Cook." On the day Remy learns his hero has died, he is evicted and ends up alone in Paris. By luck, he discovers Gasteau's restaurant, down to three stars and run by a frozen-food-hawking chef. As Remy enters, so does Linguini, a clumsy youth hired as a garbage boy. To save the soup that Linguini accidentally fouls, Remy throws in some ingredients; the soup is a success and Linguini's career as a chef is born. Can Remy find a way to maintain the fiction and use his gift?
Remy is a rat, constantly risking life in an expensive French restaurant because of his love of good food, as well as a desire to become a chef. Yet, obviously, this is a rather tough dream for a rat. But opportunity knocks when a young boy, who desperately needs to keep his job at the restaurant, despite his lack of cooking abilities, discovers and partners the young Remy. Its up to the two of them to avoid the insane head chef, bring the rest of Remy's family up to his standards, win his partner a girl, and, of course, produce the finest Ratatouille in all of France.
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